龍蝨科 (学名:Dytiscidae ),亦作龍虱科 ,為水甲蟲 的一種,隸屬於鞘翅目 之下的肉食亞目 。生活在田野、水溝、小溪等水體中,是一種掠食性水生昆蟲 。其學名源於希臘文δυτικός (dytikos ),意思就是「肉食性的潛水甲蟲」。本科物種幾乎見於全世界所有淡水棲境,但亦有部分於落葉堆中棲息[1] 。成蟲體長一般在1和2.5 cm(0.4—1.0英寸)之間,儘管物種間的差異會比較大。當中見於歐洲的Dytiscus latissimus 及巴西的Megadytes ducalis 有最大的體型,可分別長至4.5 cm(1.8英寸)到4.75 cm(1.9英寸)[1] [2] 。與之相反,擁有最小體型的物種應當為棲息於澳大利亞 地下水中的Limbodessus atypicali ,只有約0.9 mm(0.035英寸)長[1] 。大多數物種的顏色都在深棕色、黑色或深橄榄绿 ,某些亚科 的物種會在其邊沿帶有金黃色的條狀斑紋。
食用 本科物種的成蟲,特别是Cybister 屬的物種,均為可食用昆蟲 。在美國内华达州 洪堡乾湖 的一個山洞内,還在當地史前人類遺下的粪化石 内發現有Cybister explanatus 被食用的殘骸[3] 。在墨西哥 ,他們在吃C. explanatus 時會以墨西哥夹饼 夾着來吃。在日本多個地區(例如:長野縣 )有以日本龍蝨 (Cybister japonicus )為食的習慣;在中华人民共和国 广东省 ,除了日本龍蝨以外,亦會食用C. bengalensis 、C. guerini 、C. limbatus 、C. sugillatus 、C. tripunctatus 等物種。D. marginalis 很可能也包括在內,但這個物種的肉很少。也有報告指在台灣 、泰国 及新幾內亞 亦有食用本科物種的習慣[4] 。
一隻詳細分類未明的本科物種 大但在陸地上很慢,成年後也不是特別兇猛,它們也被許多中型鳥類、哺乳動物和其他大型捕食者津津有味地吃掉。幼蟲通常比較安全,由於它們的保護色和通過水柱逃跑的能力;它們可能相當難抓,可能成為小池塘的頂級掠食者。
系統分類 本科包括下列各屬[5] [6] :
Acilius Leach, 1817 Aethionectes Sharp, 1882 Africodytes Biström, 1988 Africophilus Guignot, 1948 Agabetes Crotch, 1873 Agabinus Crotch, 1873 豆龍蝨屬 Agabus Leach, 1817[6] ,端毛龙虱属 Agametrus Sharp, 1881 Agaporomorphus Zimmermann, 1921 Aglymbus Sharp, 1880 Agnoshydrus Biström, Nilsson & Wewalka, 1997 Allodessus Guignot, 1953 彩斑龍蝨屬 Allopachria Zimmermann, 1924 Amarodytes Régimbart, 1900 Amurodytes Fery & Petrov, 2013 Andex Sharp, 1882 Andonectes Guéorguiev, 1971 Anginopachria Wewalka, Balke & Hendrich, 2001 Anodocheilus Babington, 1841 Antiporus Sharp, 1882 Australphilus Watts, 1978 Austrelatus Shaverdo, Hájek, Hendrich, Surbakti, Panjaitan & Balke, 2023 Austrodytes Watts, 1978 Barrethydrus Barretthydrus Lea, 1927Barretthydrus geminatus Lea, 1927Barretthydrus stepheni Watts, 1978Barretthydrus tibialis Lea, 1927 Batrachomatus Clark, 1863 Belladessus K.B.Miller & Short, 2015 Bidessini Sharp, 1880 Bidessodes Régimbart, 1895 Bidessonotus Régimbart, 1895 Bidessus Sharp, 1882 Boreonectes (Fery & Ribera, 2018) Fery & Ribera, 2018 Boreonectes Angus, 2010 Borneodessus Balke, Hendrich, Mazzoldi & Biström, 2002 Brachyvatus Zimmermann, 1919 Brancuporus Hendrich, Toussaint & Balke, 2014 Bunites Spangler, 1972 Calicovatellus K.B.Miller & Lubkin, 2001 Canthyporus Zimmermann, 1919 Capelatus Turner & Bilton, 2015 Caperhantus Balke, Hájek & Hendrich, 2017 Carabdytes Balke, Hendrich & Wewalka, 1992 Carabhudrus Carabhydrus Watts, 1978 Carrhydrus Celina Aubé, 1837 Chostonectes Sharp, 1880 Clarkhydrus Fery & Ribera, 2018 Clemnius Villastrigo, Ribera, Manuel, Millán & Fery, 2017 Clypeodytes Régimbart, 1894 Coelambus Thomson, 1860Coelambus cribrarius (Scudder, 1900)Coelambus derelictus (Scudder, 1900)Coelambus disjectus (Scudder, 1900)Coelambus dzieduszyckii (Lomnicki, 1894)Coelambus ganglbaueri (Lomnicki, 1894)Coelambus infernalis (Scudder, 1900)Coelambus latefasciatus (Lomnicki, 1894)Coelambus miocenus (Wickham, 1912)Coelambus niedzwiedzkii (Lomnicki, 1894)Coelambus ozokeriticus (Lomnicki, 1894)Coelambus picipoides (Lomnicki, 1894) Coelhydrus Sharp, 1882 Colymbetes Clairville, 1806 Colymbetes Schellenberg, 1806 Comaldessus Spangler & Barr, 1995 Copelatus Erichson, 1832Copelatus abonnenci Guignot, 1939Copelatus acamas Guignot, 1955Copelatus adelbert Megna, Atthakor, Manaono, Hendrich & Balke, 2017Copelatus advena Sharp, 1882Copelatus aemulus Bilardo & Rocchi, 1995Copelatus aequatorius Régimbart, 1899Copelatus aethiopicus Régimbart, 1906Copelatus agaroensis Balke, 2008Copelatus agrias Guignot, 1954Copelatus alternatus Sharp, 1882Copelatus amaroides Guignot, 1952Copelatus amatolensis Omer-Cooper, 1965Copelatus amazonicus Régimbart, 1889Copelatus ambiguus Bertrand & Legros, 1975Copelatus amphibius Ranarilalatiana & Bergsten, 2019Copelatus anastomosans Guignot, 1952Copelatus andobonicus Guignot, 1960Copelatus andreinii Régimbart, 1905Copelatus andrewesi J.Balfour-Browne, 1939Copelatus angolensis Peschet, 1924Copelatus angouei Bilardo & Rocchi, 2011Copelatus angustus Gschwendtner, 1932Copelatus ankaratra Ranarilalatiana & Bergsten, 2019Copelatus annobomensis J.Balfour-Browne, 1939Copelatus anthracinus Régimbart, 1895Copelatus antoniorum Hájek & Brancucci, 2011Copelatus aphroditae Balke, 2003Copelatus apicinotatus Bilardo & Rocchi, 2018Copelatus assalyi Bilardo & Rocchi, 2010Copelatus assimilis Régimbart, 1895Copelatus ateles Guignot, 1955Copelatus atrosulcatus Régimbart, 1906Copelatus babyrousa Hájek, Hendrich & Balke, 2018Copelatus bacchusi Wewalka, 1981Copelatus bacillifer Guignot, 1955Copelatus baculiformis Guignot, 1955Copelatus badeni Sharp, 1882Copelatus bakewelli J.Balfour-Browne, 1939Copelatus baoulicus Bilardo & Pederzani, 1978Copelatus bapounouensis Bilardo & Rocchi, 2019Copelatus baranensis Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus barbouri Young, 1942Copelatus basifasciatus Régimbart, 1895Copelatus basilewskyi Bilardo & Pederzani, 1979Copelatus bechynei Guignot, 1953Copelatus befasicus Guignot, 1956Copelatus bengalensis Guignot, 1955Copelatus bertrandi Nilsson, Bilardo & Rocchi, 1996Copelatus betampona Ranarilalatiana & Bergsten, 2019Copelatus bezdeki Sheth, Ghate & Hájek, 2018Copelatus bibulus Guignot, 1948Copelatus bicolor (Fabricius, 1792) Copelatus biformis Sharp, 1882Copelatus bimaculatus (Resende & Vanin, 1991) Copelatus binaghii Bilardo & Pederzani, 1978Copelatus biolleyi Guignot, 1952Copelatus biroi Guignot, 1956Copelatus birougouensis Bilardo & Rocchi, 2018Copelatus biseriatus J.Balfour-Browne, 1939Copelatus biswasi Mukherjee & Sengupta, 1986Copelatus blancasi Guignot, 1958Copelatus blatchleyi Young, 1953Copelatus bolivianus Guignot, 1957Copelatus bombycinus Guignot, 1952Copelatus bonvouloiri Sharp, 1882Copelatus bottegoi Régimbart, 1895Copelatus bougainvillensis Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus boukali Hendrich & Balke, 1998Copelatus brancuccii Rocchi, 1979Copelatus brasiliensis Zimmermann, 1921Copelatus brendelli Hájek, Hendrich & Balke, 2018Copelatus brevicornis (Sharp, 1882) Copelatus brevistrigatus Guignot, 1959Copelatus brivioi Rocchi, 1976Copelatus bromeliarum (Scott, 1912) Copelatus brullei Aubé, 1838Copelatus brunneus J.Balfour-Browne, 1939Copelatus buqueti Aubé, 1838Copelatus burgeoni Gschwendtner, 1930Copelatus caelatipennis Aubé, 1838Copelatus caelatipennis Chevrolat, 1863Copelatus caelatus Guignot, 1952Copelatus caffer J.Balfour-Browne, 1939Copelatus calaquei Bilardo & Rocchi, 2008Copelatus camerunensis Guignot, 1941Copelatus capensis Sharp, 1882Copelatus carayoni Legros, 1949Copelatus carinatus Sharp, 1882Copelatus celinoides Guignot, 1952Copelatus cessaima Caetano, Bená & Vanin, 2013Copelatus chevrolati Aubé, 1838Copelatus chibcha Guignot, 1952Copelatus chinensis Regimbart, 1899Copelatus chipiriricus Guignot, 1957Copelatus chloroticus Régimbart, 1899Copelatus cinnamomeus Régimbart, 1895Copelatus clarki Sharp, 1882Copelatus collarti Gschwendtner, 1932Copelatus compertus Guignot, 1956Copelatus concii Bilardo, 1982Copelatus concolor Sharp, 1882Copelatus concolorans J.Balfour-Browne, 1939Copelatus confinis Bilardo & Rocchi, 1999Copelatus congo Gschwendtner, 1938Copelatus consimilis Bilardo & Rocchi, 2002Copelatus consors Sharp, 1882Copelatus cooperae Bilardo & Pederzani, 1972Copelatus cordovai Megna & Epler, 2012Copelatus cordylinoides J.Balfour-Browne, 1938Copelatus coxalis Sharp, 1882Copelatus crassus Régimbart, 1895Copelatus cryptarchoides Régimbart, 1899Copelatus cubaensis Schaeffer, 1908Copelatus curtistriatus Bilardo & Rocchi, 1995Copelatus curvispinis Bilardo & Rocchi, 2008Copelatus cynthiae Bilardo & Rocchi, 2008Copelatus daemeli Sharp, 1882Copelatus danyi Megna & Epler, 2012Copelatus darlingtoni Young, 1942Copelatus davidi Wewalka, 2017Copelatus debilis Sharp, 1882Copelatus deccanensis Sheth, Ghate & Hájek, 2018Copelatus decellei Bilardo, 1982Copelatus decemsulcatus Régimbart, 1895Copelatus deceptor Bilardo & Rocchi, 1995Copelatus dentatipenis Jiang, Hajek & Jia, 2022Copelatus depressus Sharp, 1882Copelatus descarpentriesi Bertrand & Legros, 1975Copelatus diffisus Guignot, 1939Copelatus dimorphus Sharp, 1882Copelatus distinctus Aubé, 1838Copelatus distinguendus Régimbart, 1903Copelatus diversistriatus Jiang, Hajek & Jia, 2022Copelatus djiboutensis Wewalka & Jäch, 2017Copelatus dolosus Guignot, 1956Copelatus doriae Sharp, 1882Copelatus doudouensis Bilardo & Rocchi, 2015Copelatus duodecimstriatus Aubé, 1838Copelatus duponti Aubé, 1838Copelatus efoutensis Bilardo & Rocchi, 1995Copelatus ejactus Omer-Cooper, 1965Copelatus ellai Bilardo & Rocchi, 1995Copelatus elutus Guignot, 1958Copelatus enganensis Guignot, 1940Copelatus epactus Guignot, 1948Copelatus erichsonii Guérin-Méneville, 1847Copelatus espinhasso Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024[7] Copelatus esteriensis Bilardo & Pederzani, 1978Copelatus eucritus Guignot, 1952Copelatus evanidus Bilardo & Rocchi, 1995Copelatus exaratus Sharp, 1882Copelatus externus Kirsch, 1873Copelatus falciformis Pederzani & Schizzerotto, 2017Copelatus fasciatus Bilardo & Rocchi, 1995Copelatus fastidiosus Guignot, 1959Copelatus feae Régimbart, 1888Copelatus felicis Hajek, Jiang & Jia, 2022Copelatus fernandensis Guignot, 1952Copelatus ferruginicollis Régimbart, 1895Copelatus ferus Guignot, 1953Copelatus festae Régimbart, 1899Copelatus fidschiensis Zimmermann, 1928Copelatus fijiensis Guignot, 1955Copelatus filiformis Sharp, 1882Copelatus fizpaci Bilardo & Rocchi, 2013Copelatus flavicans Guignot, 1952Copelatus flavidus Régimbart, 1895Copelatus florae Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024[7] Copelatus fluviaticus Guignot, 1957Copelatus fontanus J.Balfour-Browne, 1950Copelatus fossilis Říha, 1974 Copelatus fractistriatus Bilardo & Rocchi, 1995Copelatus freudei Guignot, 1955Copelatus fryi J.Balfour-Browne, 1939Copelatus fulviceps J.Balfour-Browne, 1938Copelatus fuscipennis Sharp, 1882Copelatus fuscomaculatus Guignot, 1952Copelatus gabonicus Bilardo & Pederzani, 1978Copelatus galapagoensis G.R.Waterhouse, 1845Copelatus garambanus Guignot, 1955Copelatus gardineri Scott, 1912Copelatus geayi Régimbart, 1904Copelatus geniculatus Sharp, 1882Copelatus gentilis Sharp, 1882Copelatus gestroi (Sharp, 1882) Copelatus gigas Bilardo & Rocchi, 2019Copelatus glyphicus (Say, 1823) Copelatus griffinii Régimbart, 1899Copelatus gschwendtneri Guignot, 1939Copelatus guadelupensis Legros, 1948Copelatus guerini Aubé, 1838Copelatus guineensis Guignot, 1955Copelatus haemorrhoidalis Régimbart, 1883Copelatus hararensis Guignot, 1952Copelatus hardenbergi J.Balfour-Browne, 1950Copelatus hebeter Guignot, 1953Copelatus heterogynus Régimbart, 1899Copelatus hydroporoides (Murray, 1859) Copelatus ibrahimi Zalat, Saleh, Angus & Kaschef, 2000Copelatus iguelaensis Bilardo & Rocchi, 2002Copelatus ilybioides Régimbart, 1895Copelatus imitator Bilardo & Rocchi, 2002Copelatus inaequalis Sharp, 1882Copelatus incognitus Sharp, 1882Copelatus indicus Sharp, 1882Copelatus inopinatus Bilardo & Rocchi, 1995Copelatus inornatus Sharp, 1882Copelatus insidiosus Bilardo & Rocchi, 1995Copelatus insolitus Chevrolat, 1863Copelatus instabilis Régimbart, 1897Copelatus instriatus Zimmermann, 1921Copelatus insuetus Guignot, 1941Copelatus insulanus Guignot, 1939Copelatus integer Sharp, 1882Copelatus internus Régimbart, 1904Copelatus interstriatus J.Balfour-Browne, 1939Copelatus inuber Guignot, 1952Copelatus ipiformis Régimbart, 1895Copelatus irinus Régimbart, 1899Copelatus irregularis W.J.MacLeay, 1871Copelatus ischius Guignot, 1956Copelatus jactator Guignot, 1955Copelatus jaechi Bilardo & Rocchi, 2016Copelatus jamaicensis Young, 1942Copelatus japonicus Sharp, 1884Copelatus jarrigei Legros, 1954Copelatus javanus Régimbart, 1883Copelatus jocosus Guignot, 1955Copelatus johannis Guignot, 1954Copelatus kalaharii Gschwendtner, 1935Copelatus kammuriensis Tamu & Tsukamoto, 1955Copelatus kaszabi Guignot, 1956Copelatus kely Ranarilalatiana & Bergsten, 2019Copelatus kietensis Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus kilimandjarensis Bilardo & Pederzani, 1972Copelatus kindianus Guignot, 1954Copelatus kongouensis Bilardo & Rocchi, 1999Copelatus koreanus Mori, 1932Copelatus laccophilinus Sharp, 1882Copelatus laeticulus Sharp, 1882Copelatus laevipennis Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus lamottei Legros, 1954Copelatus lanzai Bilardo & Rocchi, 1995Copelatus laraensis Bilardo & Rocchi, 1995Copelatus lasckonyi Bilardo & Rocchi, 1995Copelatus latens Guignot, 1952Copelatus laticollis Régimbart, 1899Copelatus latifasciatus Bilardo & Rocchi, 1999Copelatus latipes Sharp, 1882Copelatus latus J.Balfour-Browne, 1950Copelatus leonardii Bilardo & Rocchi, 1999Copelatus leonensis Legros, 1954Copelatus lepersonneae Gschwendtner, 1943Copelatus lignosus Guignot, 1955Copelatus lineatipennis Guignot, 1955Copelatus lineatus (Guérin-Méneville, 1838) Copelatus longicornis Sharp, 1882Copelatus lootensi Guignot, 1955Copelatus louayensis Bilardo & Rocchi, 2004Copelatus luctuosus Guignot, 1939Copelatus luridescens Régimbart, 1889Copelatus luteocinctus Guignot, 1955Copelatus luteomaculatus Guignot, 1956Copelatus luzonicus Guignot, 1952Copelatus macellus Guignot, 1950Copelatus madoni Guignot, 1952Copelatus mahajanga Pederzani & Hájek, 2005Copelatus mahleri Holmen & Vazirani, 1990Copelatus makokouensis Bilardo & Rocchi, 1995Copelatus malaisei Guignot, 1954Copelatus malavergnorum Manuel & Ramahandrison, 2022Copelatus mancus Sharp, 1887Copelatus marginatus Sharp, 1882Copelatus marginipennis (Laporte, 1835) Copelatus martinbaehri Hendrich, Shaverdo, Hájek & Balke, 2019Copelatus martini Manuel, Deler-Hernández, Megna & Hájek, 2018Copelatus masculinus Régimbart, 1899Copelatus massaicus Guignot, 1941Copelatus mathani Guignot, 1952Copelatus maushomi Sheth, Ghate & Hájek, 2018Copelatus mbokoensis Bilardo & Rocchi, 2006Copelatus melanogrammus Régimbart, 1883Copelatus mendax Bilardo & Rocchi, 2011Copelatus miaowangi Hajek, Jiang & Jia, 2022Copelatus minimus Bilardo & Rocchi, 1995Copelatus minor Bilardo & Pederzani, 1978Copelatus minutissimus J.Balfour-Browne, 1939Copelatus mocquerysi Régimbart, 1895Copelatus mohelicus Guignot, 1959Copelatus monticola Guignot, 1951Copelatus montivagus Young, 1942Copelatus mopanshanensis Jiang, Zhao & Hajek, 2022Copelatus mulangensis Bameul, 2003Copelatus mundus Sharp, 1882Copelatus mutabilis Bilardo & Rocchi, 1999Copelatus mvoungensis Bilardo & Rocchi, 2004Copelatus mysorensis Vazirani, 1970Copelatus nakamurai Guéorguiev, 1970Copelatus nangaensis Guignot, 1952Copelatus nanlingensis Hajek, Jiang & Jia, 2022Copelatus nauclerus Guignot, 1939Copelatus neavei J.Balfour-Browne, 1950Copelatus neelumae Vazirani, 1973Copelatus neogestroi Balke, 2008Copelatus neoguineensis Zimmermann, 1919Copelatus nguemai Bilardo & Rocchi, 2011Copelatus nigricans Sharp, 1882Copelatus nigricollis Paulino de Oliveira, 1880 Copelatus nigrolineatus Sharp, 1882Copelatus nigropennis Zimmermann, 1927Copelatus nigrostriatus Régimbart, 1895Copelatus nilssoni Wewalka, 1982Copelatus nimbaensis Legros, 1958Copelatus nitens Bilardo & Rocchi, 1999Copelatus nitidus Sharp, 1882Copelatus nodieri Régimbart, 1895Copelatus normalis Erichson, 1847Copelatus notius Omer-Cooper, 1965Copelatus nzei Bilardo & Rocchi, 1999Copelatus oblitus Sharp, 1882Copelatus obscurus Sharp, 1882Copelatus occultus Bilardo & Rocchi, 1995Copelatus ogasawarensis Kamiya, 1932Copelatus onorei Pederzani & Rocchi, 1982Copelatus ornatipennis Zimmermann, 1926Copelatus owas Régimbart, 1895Copelatus pachys Guignot, 1955Copelatus pallidus Régimbart, 1895Copelatus paludorum Guignot, 1952Copelatus pandanorum Scott, 1912Copelatus panguana Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024[7] Copelatus pantosi Guignot, 1958Copelatus papuensis J.Balfour-Browne, 1939Copelatus parabaptus Guignot, 1955Copelatus parallelipipedus Régimbart, 1895Copelatus parallelus Zimmermann, 1920Copelatus pardii Rocchi, 1990Copelatus parisii Guignot, 1959Copelatus parumstriatus Gschwendtner, 1934Copelatus paryphes Guignot, 1955Copelatus pederzanii Bilardo & Rocchi, 1995Copelatus pereirai Guignot, 1955Copelatus peridinus Guignot, 1955Copelatus pinnifer Guignot, 1952Copelatus piriensis Omer-Cooper, 1965Copelatus platynotus Régimbart, 1895Copelatus polystrigus Sharp, 1882Copelatus ponomarenkoi Říha, 1974 Copelatus portior Guignot, 1956Copelatus posticatus (Fabricius, 1801) Copelatus poungai Bilardo & Rocchi, 2006Copelatus predaveterus Miller, 2003Copelatus prolixus Sharp, 1882Copelatus prolongatus Sharp, 1882Copelatus propino Guignot, 1960Copelatus propinquus Régimbart, 1895Copelatus proximus Sharp, 1882Copelatus pseudostriatus Ranarilalatiana & Bergsten, 2019Copelatus pulchellus (Klug, 1834) Copelatus pulicarius Régimbart, 1895Copelatus pumilus Régimbart, 1895Copelatus punctatus Bilardo & Rocchi, 1995Copelatus punctulatus Aubé, 1838Copelatus puzhelongi Jiang, Hajek & Jia, 2022Copelatus quadrisignatus Régimbart, 1877Copelatus quadristriatus Pederzani & Schizzerotto, 2017Copelatus racenisi Guignot, 1951Copelatus ragazzii Régimbart, 1887Copelatus regimbarti Branden, 1885Copelatus restrictus Sharp, 1882Copelatus rimosus Guignot, 1952Copelatus rivalis Guignot, 1952Copelatus rocchii Bilardo, 1982Copelatus romani Zimmermann, 1924Copelatus rosulae Hajek, Jiang & Jia, 2022Copelatus royi Legros, 1958Copelatus rubiginosus Guignot, 1954Copelatus ruficapillus Régimbart, 1895Copelatus ruteri Legros, 1954Copelatus saegeri Guignot, 1955Copelatus safiotra Ranarilalatiana & Bergsten, 2019Copelatus sahlbergi J.Balfour-Browne, 1939Copelatus sallaei Sharp, 1882Copelatus sanfilippoi Bilardo & Pederzani, 1972Copelatus saverii Pederzani & Schizzerotto, 2017Copelatus scalptus Guignot, 1954Copelatus scaphites Guignot, 1955Copelatus schaefferi Young, 1942Copelatus schereri Wewalka, 1981Copelatus schuhi Hendrich & Balke, 1998Copelatus scutatus Guignot, 1952Copelatus scytalotus Guignot, 1956Copelatus senegalensis Lagar & Hernando, 1991Copelatus sexstriatus Sharp, 1882Copelatus sharpi Branden, 1885Copelatus sibelaemontis Hájek, Hendrich, Hawlitschek & Balke, 2010Copelatus silvestrii Régimbart, 1903Copelatus simoni Régimbart, 1889Copelatus singularis Bilardo & Rocchi, 1995Copelatus sociennus J.Balfour-Browne, 1952Copelatus solitarius Sharp, 1882Copelatus sordidipennis Régimbart, 1895Copelatus speciosus Régimbart, 1892Copelatus spoliatus Guignot, 1955Copelatus stavropolitanus Říha, 1974 Copelatus stillicidicola Bilardo & Rocchi, 1995Copelatus striatellus Boheman, 1848Copelatus striaticollis Lucas, 1857Copelatus striatopterus Aubé, 1838Copelatus striatulus Aubé, 1838Copelatus strigipennis (Laporte, 1835) Copelatus strigosulus (Fairmaire, 1878) Copelatus strinatii Guignot, 1958Copelatus striolatus Peschet, 1917Copelatus stygis Guignot, 1958Copelatus subdeficiens Régimbart, 1902Copelatus subsimilis Guignot, 1958Copelatus substriatus Kirsch, 1873Copelatus subterraneus Guéorguiev, 1978Copelatus sudrei Bameul, 2003Copelatus sulcatus Sharp, 1882Copelatus sulcipennis (Laporte, 1835) Copelatus sumbawensis Régimbart, 1899Copelatus suppar Guignot, 1956Copelatus supplementaris Régimbart, 1895Copelatus sylvaticoides Bilardo & Rocchi, 2013Copelatus sylvaticus Guignot, 1952Copelatus takakurai Satô, 1985Copelatus tanaus Guignot, 1953Copelatus taprobanicus Wewalka & Vazirani, 1985Copelatus tenebrosis Regimbart, 1880Copelatus tenebrosus Régimbart, 1880Copelatus tengchongensis Hajek, Jiang & Jia, 2022Copelatus teranishii Kamiya, 1938Copelatus terminalis Sharp, 1882Copelatus ternatensis Régimbart, 1899Copelatus terzanii Bilardo & Saverio, 2016Copelatus thiriouxi Peschet, 1917Copelatus thrasys Guignot, 1952Copelatus tibialis Sharp, 1882Copelatus tinctor Guignot, 1954Copelatus togoensis Régimbart, 1895Copelatus tomokunii Satô, 1985Copelatus tondangoyei Bilardo & Rocchi, 2013Copelatus tostus J.Balfour-Browne, 1950Copelatus trifilis Guignot, 1958Copelatus triglyphus Guignot, 1955Copelatus trilobatus Régimbart, 1895Copelatus tschaga Bilardo & Pederzani, 1972Copelatus tucuchiensis J.Balfour-Browne, 1939Copelatus tulagicus Guignot, 1942Copelatus uludanuensis Hendrich & Balke, 1995Copelatus undecimstriatus Aubé, 1838Copelatus urceolus Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus usagarensis Zimmermann, 1926Copelatus vagatus Guignot, 1952Copelatus vagestriatus Zimmermann, 1919Copelatus vagus Bilardo & Rocchi, 1995Copelatus validus Sharp, 1882Copelatus vancli Hájek, Hendrich & Balke, 2018Copelatus vanninii Bilardo & Rocchi, 1999Copelatus variegatus Régimbart, 1895Copelatus variistriatus Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus venustus Bilardo & Rocchi, 1995Copelatus vigintistriatus Fairmaire, 1869Copelatus vigintisulcatus Régimbart, 1895Copelatus villiersi Guignot, 1950Copelatus virungaensis Bilardo, 1982Copelatus vitraci Legros, 1948Copelatus vivax Guignot, 1953Copelatus vokoka Ranarilalatiana & Bergsten, 2019Copelatus wallacei J.Balfour-Browne, 1939Copelatus waltoni J.Balfour-Browne, 1950Copelatus wayanadensis Manivannan & Madani, 2011Copelatus wewalkai Holmen & Vazirani, 1990Copelatus weymarni J.Balfour-Browne, 1947Copelatus xanthocephalus Régimbart, 1899Copelatus xiniudong Jiang, Hajek & Jia, 2022Copelatus yacumensis Guignot, 1957Copelatus yaguarete Scheers, 2018Copelatus yombensis Bilardo & Rocchi, 2015Copelatus zadiensis Bilardo & Rocchi, 1995Copelatus zanabato Manuel & Ramahandrison, 2022Copelatus zanatanensis Ranarilalatiana & Bergsten, 2019Copelatus zimmermanni Gschwendtner, 1934 Copelatus Sheth, Ghate & Hájek, 2018 Coptotomus Say, 1830 Cretodytes Ponomarenko, 1977 Crinodessus K.B.Miller, 1997 大龍蝨屬 Cybister Curtis, 1827,真龙虱属 Darwinhydrus Sharp, 1882 Deronectes Sharp, 1882 Derovatellus Sharp, 1882 Desmopachria Babington, 1841 Deuteronectes Guignot, 1945 Deuteronectus Dimitshydrus Uéno, 1996 Dyschirioides Dyticus Geoffroy, 1762 Dytiscus Linnaeus, 1758 Electruphilus Balker, Toledo & Hendrich, 2019 Ereboporus K.B.Miller, Gibson & Alarie, 2009 Eretes Laporte, 1833 Etruscodytes Mazza, Cianferoni & Rocchi, 2013 Exocelina Broun, 1886 Fontidessus K.B.Miller & Spangler, 2008 Geodessus Brancucci, 1979 Gibbidessus Watts, 1978 Glareadessus Wewalka & Biström, 1998 Graphoderus Dejean, 1833 Graptodytes Seidlitz, 1887 Haideoporus Young & Longley, 1976 Hamadiana Benetti, Short & Michat, 2019 Hemibidessus Zimmermann, 1921 Heroceras Guignot, 1950 Heterhydrus Fairmaire, 1869 Heterosternuta Strand, 1935 Hintonia Young, 1980 Hoperius Fall, 1927 Hornectes Fery & Ribera, 2018 Hovahydrus Biström, 1982 Huxelhydrus Sharp, 1882 Hydaticus Leach, 1817 Hyderodes Hope, 1838 Hydrocolus Roughley & Larson, 2000 Hydrodessus J.Balfour-Browne, 1953 Hydrodytes K.B.Miller, 2001 短褶龙虱属 Hydroglyphus Motschulsky, 1853 Hydronebrius Jakovlev, 1897 Hydropeplus Sharp, 1882 Hydroporus Clairville, 1806 Hydroporus Schellenberg, 1806 Hydrotrupes Sharp, 1882 Hydrovatus Motschulsky, 1853 水龙虱属 Hygrotus Stephens, 1828 Hyphoporus Sharp, 1882 Hyphovatus Wewalka & Biström, 1994 异爪龙虱属 Hyphydrus Illiger, 1802 Hypodessus Guignot, 1939 Iberonectes Fery & Ribera, 2018 Iberoporus Castro & Delgado, 2001 Ilybiosoma Crotch, 1873 Ilybius Erichson, 1832 Incomptodessus K.B.Miller & García, 2011 Isonotus Houlbert, 1934 Japanolaccophilus Satô, 1972 Kakadudessus Hendrich & Balke, 2009Kakadudessus tomweiri Hendrich & Balke, 2009 Kuschelydrus Ordish, 1976 Laccodytes Régimbart, 1895 Laccomimus Toledo & Michat, 2015 Lacconectus Motschulsky, 1855 粒龙虱属 Laccophilus Leach, 1815 Laccoporus J.Balfour-Browne, 1939Laccoporus nigritulus (Gschwendtner, 1936) Laccornellus Roughley & Wolfe, 1987 Laccornis Gozis, 1914 Laccosternus Brancucci, 1983 Lancetes Sharp, 1882 Laodytes Queney, Lemaire & Ferrand, 2020 Larsonectes Fery & Ribera, 2018 Leconectes Fery & Ribera, 2018 Leiodytes Guignot, 1936 Leuronectes Sharp, 1882 Liadyxianus Prokin, Petrov, Wang & Ponomarenko, 2013 Limbodessus Guignot, 1939 Liodessus Guignot, 1939 Lioporeus Guignot, 1950 Liopterus Dejean, 1833 Madaglymbus Shaverdo & Balke, 2008 Matus Aubé, 1836 Megadytes Sharp, 1882 Megaporus Brinck, 1943 Meladema Laporte, 1835 Melanodytes Seidlitz, 1887 Meridiorhantus Balke, Hájek & Hendrich, 2017 Mesodytes Prokin, Petrov, Wang & Ponomarenko, 2013 Metaporus Guignot, 1945 Methles Sharp, 1882 Microdessus Young, 1967 Microdytes J.Balfour-Browne, 1946 Microhydrodytes K.B.Miller, 2002 Miodytiscus Wickham, 1911 Morimotoa Uéno, 1957 Mystonectes Fery & Ribera, 2018 Napodytes Steiner, 1981 Nartus Zaitzev, 1907 孔龙虱属 Nebrioporus Régimbart, 1906 Necterosoma Sharp, 1882 Necterosoma W.J.MacLeay, 1871 Nectoboreus Fery & Ribera, 2018 Nectomimus Fery & Ribera, 2018 Nectoporus Guignot, 1950 Neobidessodes Hendrich & Balke, 2009 Neobidessus Young, 1967 Neoclypeodytes Young, 1967 Neonectes J.Balfour-Browne, 1944 Neonectes Zimmermann, 1932 Neoporus Guignot, 1931 Neoscutopterus J.Balfour-Browne, 1943 彩裳粒龙虱属 Neptosternus Sharp, 1882 Notaticus Zimmermann, 1928 Novadessus K.B.Miller, 2016 Onychohydrus Schaum & White, 1847 Oreodytes Seidlitz, 1887 Pachydrus Sharp, 1882 Pachynectes Régimbart, 1903 Palaeodytes Ponomarenko, 1987 Papuadessus Balke, 2001 Paroster Sharp, 1882 Peschetius Guignot, 1942 Petrodessus K.B.Miller, 2012 Philaccolilus Guignot, 1937 Philaccolus Guignot, 1937 Philodytes J.Balfour-Browne, 1939 Phreatodessus Ordish, 1976 Platambus Thomson, 1859 Platydytes Biström, 1988 Platynectes Gueorguiev, 1972 Platynectes Régimbart, 1879 Porhydrus Guignot, 1945 Primospes Sharp, 1882 Procoelambus Théobald, 1937 Prodaticus Sharp, 1882 缝龙虱属 Pseuduvarus Biström, 1988 Psychopomporus Jean, Telles & K.B.Miller, 2012 Pteroporus Guignot, 1933 Queda Sharp, 1882 Regimbartina Chatanay, 1911 Rhantaticus Sharp, 1880 Rhantus Dejean, 1833 Rhithrodytes Bameul, 1989 Rompindessus Balke, Bergsten & Hendrich, 2017 Sandracottus Sharp, 1882 Sanfilippodytes Franciscolo, 1979 Scarodytes Gozis, 1914 Schistomerus Palmer, 1957 Sekaliporus Watts, 1997 Sharphydrus Omer-Cooper, 1958 Siamoporus Spangler, 1996 Siettitia Abeille de Perrin, 1904 Sinodytes Spangler, 1996 Spanglerodessus K.B.Miller & García, 2011 Spencerhydrus Sharp, 1882 Sternhydrus Brinck, 1945 Sternopriscus Sharp, 1880 Stictonectes Brinck, 1943 Stictotarsus Zimmermann, 1919 Stygoporus Larson & LaBonte, 1994 Suphisellus Zimmermann, 1921 Tassilodytes Fery & Bouzid, 2016 Tepuidessus Spangler, 1981 Thermonectus Dejean, 1833Thermonectus alfredi Griffini, 1898Thermonectus basillaris (Harris, 1829)Thermonectus batesi Sharp, 1882Thermonectus circumscriptus (Latreille, 1809)Thermonectus cuneatus Sharp, 1882 Thermonectus depictus Sharp, 1882 Thermonectus duponti (Aubé, 1838)Thermonectus intermedius Crotch, 1873 Thermonectus laporti (Aubé, 1838)Thermonectus leprieuri J.Balfour-Browne, 1944Thermonectus margineguttatus (Aubé, 1838)Thermonectus marmoratus (Gray, 1831)Thermonectus nigrofasciatus (Aubé, 1838)Thermonectus nobilis Zimmermann, 1924Thermonectus sibleyi Goodhue-McWilliams, 1981Thermonectus simulator Sharp, 1882Thermonectus succinctus (Aubé, 1838)Thermonectus tremouillesi Michat & Torres, 2016[8] Thermonectus variegatus (Dejean, 1836)Thermonectus variegatus (Laporte de Castelnau, 1835)Thermonectus zimmermani Goodhue-McWilliams, 1981 Tikoloshanes Omer-Cooper, 1956 Tiporus Watts, 1985 Trichonectes Guignot, 1941 Trogloguignotus Sanfilippo, 1958 Tyndallhydrus Sharp, 1882 Typhlodessus Brancucci, 1985 Uvarus Guignot, 1939 Vatellus Aubé, 1837 Yola Gozis, 1886 Yolina Guignot, 1936 Zaitzevhydrus Fery & Ribera, 2018 Zimpherus K.B.Miller & Wheeler, 2015 參考文獻 联邦科学与工业研究组织 (CSIRO) (2004): Water for a Healthy Country - Family Dytiscidae (页面存档备份 ,存于互联网档案馆 ). Version of 2004-JUL-02. Retrieved 2008-AUG-04De Foliart, Gene R. (2002): Chapter 26 - Eastern Asia: China, Japan, and other countries . In: The Human Use of Insects as a Food Resource: A Bibliographic Account in Progress .Jäch, Manfred A. (2003): Fried water beetles Cantonese style. American Entomologist 49 (1): 34-37. PDF fulltext Larson, D.J., Alarie, Y., and Roughley, R.E. (2000): Predaceous Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa. ISBN 978-0-660-17967-4 .外部連結
溝胸長扁蟲科 長扁甲科 侏羅甲科 微長扁蟲科 澳州長扁蟲科
長蠹蟲下目
扁蟲下目
金花蟲總科 郭公蟲總科 扁蟲總科 擬瓢甲科 毛蕈蟲科 澳洲蕈蟲科 長蕈蟲科 擬吸木蟲科 洞颚朽木甲科 朽葉蟲科 瓢蟲科 瑣微蟲科 吸木蟲科 扁蟲科 Cyclaxyridae 盤甲科 伪瓢虫科 大蕈蟲科 蜡斑甲科 Hobartiidae 小出尾虫科 姬扁甲科 Lamingtoniidae (Lamingtonium binnaberrense ) 姬薪蟲科 出尾扁甲科 Myraboliidae 露尾甲科 捕蠹甲科 姬花甲科 皮扁甲科 皮跳甲科 原扁甲科 細扁甲科 短甲科 姬蕈甲科 象鼻蟲總科 筒蠹蟲總科 擬步總科
叩頭蟲下目
長蠢總科 丸甲總科 花甲總科 叩頭蟲總科 Anischiidae 偽長花蚤科 頸螢科 菊虎科 樹叩甲科 Drilidae 叩頭蟲科 隱唇叩甲科 萤科 紅螢科 Omalisidae Omethidae (false fireflies) Phengodidae (glowworm beetles, long-lipped beetles) Plastoceridae (Plastocerus angulosus ) Podabrocephalidae Rhagophthalmidae Rhinorhipidae (Rhinorhipus tamborinensis ) Throscidae (false metallic wood-boring beetles) 圓花蚤總科 Clambidae Decliniidae (Declinia relicta ) Eucinetidae (plate-thigh beetles) 圓花蚤科
金龜子下目
金龜子總科 Belohinidae (Belohina inexpectata ) Bolboceratidae Ceratocanthidae Diphyllostomatidae (false stag beetles) 糞金龜科 絨毛金龜科 Glaresidae (enigmatic scarab beetles) 駝金龜科 鍬形蟲科 紅金龜科 黑蜣科 Pleocomidae (rain beetles) 金龜子科 皮金龜科
隱翅蟲亞目
閻蟲總科 牙蟲總科 Epimetopidae Georissidae Helophoridae Hydrochidae 牙蟲科 Spercheidae 隱翅蟲總科 Agyrtidae (primitive carrion beetles) 平唇水龟甲科 球蕈甲科 Ptiliidae (feather-winged beetles) Scydmaenidae (ant-like stone beetles) 葬甲科 隱翅蟲科
鞘翅目子類群列表