List of penguins

Penguins are birds in the family Spheniscidae in the monotypic order Sphenisciformes.[1] They inhabit high-productivity marine habitats, almost exclusively in the Southern Hemisphere; the only species to occur north of the Equator is the Galapagos penguin.[2][3] The only group of birds other than the ratites to be entirely flightless, penguins are extremely adapted to their aquatic lifestyle, with a streamlined shape that minimizes drag, countershaded dark-and-white plumage,[3] dense bones,[2] powerful flippers, and insulating feathers that allow them to withstand very low temperatures on land and in water.[3]

Chinstrap penguin

There are currently 18 extant species of penguins recognised by the International Ornithologists' Union, distributed among six genera.[1] Many species of fossil penguins are known from the Paleocene onwards; however, their exact number and taxonomy are unsettled due to ongoing discoveries.[4]

Conventions

IUCN Red List categories
Conservation status
 EX Extinct (0 species)
 EW Extinct in the wild (0 species)
 CR Critically Endangered (0 species)
 EN Endangered (5 species)
 VU Vulnerable (4 species)
 NT Near threatened (2 species)
 LC Least concern (7 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the penguin's range is provided. Ranges are based on the IOC World Bird List for that species unless otherwise noted. Population estimates are of the number of mature individuals and are taken from the IUCN Red List.

This list follows the taxonomic treatment (designation and order of species) and nomenclature (scientific and common names) of version 13.2 of the IOC World Bird List.[1] Where the taxonomy proposed by the IOC World Bird List conflicts with the taxonomy followed by the IUCN[a] or the 2023 edition of The Clements Checklist of Birds of the World,[6] the disagreement is noted next to the species's common name (for nomenclatural disagreements) or scientific name (for taxonomic disagreements).

Classification

The International Ornithologists' Union (IOU) recognises 18 species of penguins in six genera. This list does not include hybrid species, extinct prehistoric species, or putative species not yet accepted by the IOU.

Family Spheniscidae

Penguins

Genus AptenodytesMiller, J. F., 1778 – 2 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
King penguin

A. patagonicus
Miller, J. F., 1778
Subantarctic islands
 LC 


1,084,320–1,228,320 breeding pairs [7]

Emperor penguin

A. forsteri
Gray, G. R., 1884
Antarctic sea ice
 NT 


256,500 breeding pairs [8]

Genus PygoscelisWagler, 1832 – 3 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Adélie penguin

P. adeliae
(Hombron and Jacquinot, 1841)
Antarctica and surrounding islands
 LC 


1,084,320–1,228,320 breeding pairs [9]

Chinstrap penguin

P. antarcticus
(Forster, J. R., 1781)
Antarctic Peninsula and Balleny Islands
 LC 


8,000,000 [10]

Gentoo penguin

P. papua
(Forster, J. R., 1781)

Four subspecies
  • P. a. taeniata
  • P. a. papua
  • P. a. ellsworthi
  • P. a. poncetii
Subantarctic islands and locally in Antarctica
 LC 


774,000 [11]

Genus EudyptulaBonaparte, 1856 – 1 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Little penguin

E. minor
(Forster, J. R., 1781)

Six subspecies
  • E. a. novaehollandiae
  • E. a. iredalei
  • E. a. variabilis
  • E. a. minor
  • E. a. albosignata
  • E. a. chathamensis
Australia and New Zealand
 LC 


469,760 [12]

Genus SpheniscusBrisson, 1760 – 4 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Galapagos penguin

S. mendiculus
Sundevall, 1871
Galápagos Islands
 EN 


1,200 [13]

Humboldt penguin

S. humboldti
Meyen, 1834
South America
 VU 


23,800 [14]

Magellanic penguin

S. magellanicus
(Forster, J. R., 1781)
South America
 LC 


2,200,000–3,200,000 [15]

African penguin

S. demersus
(Linnaeus, 1758)
Coasts of Southern Africa
 EN 


41,700 [16]

Genus MegadyptesMilne-Edwards, 1880 – 1 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Yellow-eyed penguin

M. antipodes
(Hombron and Jacquinot, 1841)
New Zealand
 EN 


2,600–3,000 [17]

Genus EudyptesVieillot, 1816 – 7 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Macaroni penguin

E. chrysolophus
(Brandt, J. F., 1837)
Antarctic Peninsula, South America, and subantarctic islands in South Atlantic and Indian Oceans
 VU 


6,300,000 breeding pairs [18]

Royal penguin

E. schlegeli
Finsch, 1876
Macquarie Island and nearby islands LC 


1,340,000–1,660,000 [19]

Northern rockhopper penguin

E. moseleyi
Mathews & Iredale, 1921
Tristan da Cunha, Amsterdam Island, and St. Paul Island EN 


413,700 [20]

Southern rockhopper penguin

E. chrysocome
(Forster, J. R., 1781)

Two subspecies
  • E. c. filholi
  • E. c. chrysocome
South America and subantarctic islands in the Indian Ocean and New Zealand VU 


2,500,000 [21]

Fiordland penguin

E. pachyrhynchus
Gray, G. R., 1845
New Zealand
 NT 


12,500–50,000 [22]

Snares penguin

E. robustus
Oliver, 1953
Snares Islands
 VU 


63,000 [23]

Erect-crested penguin

E. sclateri
Buller, 1888
Bounty and Antipodes Islands
 EN 


150,000 [24]

Fossil genera

Basal Sphenisciformes

SpeciesNotesSource
Waimanu manneringiSlack et al. 2006
Muriwaimanu tuatahiMayr et al. 2017
Sequiwaimanu roseaeMayr et al. 2017
Crossvallia unienwilliaJadwiszczak et al. 2013
?Crossvallia waiparensisMayr et al. 2019
Petradyptes stonehouseiKsepka et al. 2023
Kaiika maxwelliFordyce & Thoman 2011
Kumimanu biceaeMayr et al. 2017
Kumimanu fordecyiKsepka et al. 2023
Kupoupou stilwelliBlokland et al. 2019

Advanced Sphenisciformes

SpeciesNotesSource
Anthropornis nordenskjoldiiJadwiszczak et al. 2002
Anthropornis grandisJadwiszczak et al. 2002
UCMP 321021 (Anthropornis sp.)Ksepka & Clarke, 2010
Palaeeudyptes antarcticus
Palaeeudyptes marplesi
Palaeeudyptes klekowskiiJadwiszczak et al. 2002
Palaeeudyptes gunnariMay represent the opposite sex of P. klekowskii [1]Jadwiszczak et al. 2002
Chilean Palaeeudyptes
Burnside "Palaeeudyptes"Ksepka & Clarke, 2010
Duntroon "Palaeeudyptes"Ksepka & Clarke, 2010
Archaeospheniscus loweiGiovanardi et al. 2021
Archaeospheniscus lopdelliGiovanardi et al. 2021
Notodyptes wimaniFormerly ArchaeospheniscusGiovanardi et al. 2021
Delphinornis larseniJadwiszczak et al. 2002
Delphinornis gracilisJadwiszczak et al. 2002
Delphinornis arctowskiiJadwiszczak et al. 2002
Mesetaornis polarisJadwiszczak et al. 2002
Marambiornis exilisJadwiszczak et al. 2002
Aprosdokitos mikroteroHospitaleche et al. 2017
Perudyptes devriesiKsepka & Clarke, 2010
Eretiscus tonniHospitaleche & Tambussi, 2008
Palaeospheniscus patagonicusHospitaleche & Tambussi, 2008
Palaeospheniscus bilocularHospitaleche & Tambussi, 2008
Palaeospheniscus bergiHospitaleche & Tambussi, 2008
Paraptenodytes antarcticusHospitaleche & Tambussi, 2008
Arthrodytes andrewsiHospitaleche & Tambussi, 2008
Madrynornis mirandus[25]Hospitaleche & Tambussi, 2008
Pachydyptes simpsoniPark & Fitzgerald, 2012
Pachydyptes ponderosusGiovanardi et al. 2021
Anthropodyptes gilliPark & Fitzgerald, 2012
Pseudaptenodytes macraeiPark & Fitzgerald, 2012
Marambiornopsis sobraliJadwiszczak et al. 2021
Icadyptes salasiClarke et al. 2007
Kairuku waewaeroaGiovanardi et al. 2021
Kairuku grebneffiGiovanardi et al. 2021
Kairuku waitakiGiovanardi et al. 2021
Glenn Murray KairukuGiovanardi et al. 2021
Platydyptes novaezealandiaeGiovanardi et al. 2021
Platydyptes marplesiGiovanardi et al. 2021
Inkayacu paracasensisClarke et al. 2010

Extinct species of extant genera

SpeciesNotesSource
Spheniscus megaramphusHospitaleche & Tambussi, 2008
Spheniscus urbinaiHospitaleche & Tambussi, 2008
Spheniscus chilensisHospitaleche & Tambussi, 2008
Spheniscus muizoniHospitaleche & Tambussi, 2008
Pygoscelis grandisHospitaleche & Tambussi, 2008
Pygoscelis tyreei
Pygoscelis calderensisHospitaleche & Tambussi, 2008
Eudyptes atatuThomas et al. 2020
Eudyptes warhamiCole et al. 2019
Eudyptes calauinaHoffmeister et al. 2014
Eudyptula wilsonaeThomas et al. 2023
Megadyptes antipodes richdaleiCole et al. 2019
Megadyptes antipodes waitahaCole et al. 2019
Aptenodytes ridgeni

Poorly understood taxa

SpeciesNotesSource
Dege hendeyi
Inguza predemersus
Duntroonornis parvus
Nucleornis insolitus
Marplesornis novaezealandiae
Korora oliveri
Paraptenodytes robustus
Platydyptes amiesi
"Pakudyptes hakataramea"Also called the "Hakataramea Penguin"Ando, 2007 (unpublished)

Invalid taxa

SpeciesNotesSource
Tonniornis mesetaensisNamed on an isolated humerus which is unable to be compared to other species present in the same locality.Ksepka & Clarke, 2010
Tonniornis minimumNamed on an isolated humerus which is unable to be compared to other species present in the same locality. Similar size to a specimen referred to Delphinornis larseni.Ksepka & Clarke, 2010
Tereingaornis moisleyiGenus is a nomen dubium due to lack of identifiable traits, but the specimen may belong to a new species (the Te Ringa Falls Penguin)Thomas et al. 2019
Wimanornis seymourensisSynonymous with Palaeeudypytes gunnari
Orthopteryx gigasHospitaleche and Reguero, 2010
Ichtyopteryx gracilisHospitaleche and Reguero, 2010
Palaeoapterodytes ictusHospitaleche, 2010
Psuedaptenodytes minorLack of diagnostic characteristics on the holotypePark & Fitzgerald, 2012

The Early Oligocene genus Cruschedula was formerly thought to belong to Spheniscidae; however, re-examination of the holotype in 1943 resulted in the genus being placed in Accipitridae.[26] Further examination in 1980 resulted in placement as Aves incertae sedis.[27]

Notes

References