Crocidurinae is a subfamily of small mammals in the shrew family Soricidae, which in turn is part of the order Eulipotyphla. A member of this family is called a crocidurine, or a white-toothed shrew. Crocidurinae is one of three subfamilies in Soricidae, along with the African shrews of Myosoricinae and the red-toothed shrews of Soricinae. They are found in Africa, Europe, and Asia, primarily in forests, savannas, shrublands, and grasslands, though some species can also be found in wetlands, rocky areas, or coastal areas. They range in size from the Etruscan shrew, one of the smallest mammal species at 3 cm (1 in) plus a 2 cm (1 in) tail,[1] to the hero shrew and Thor's hero shrew, at 15 cm (6 in) plus a 10 cm (4 in) tail. Crocidurines primarily eat invertebrates, small mammals, frogs, toads, and lizards. Almost no crocidurines have population estimates, but 21 species are categorized as endangered species, while the Andaman shrew, Christmas Island shrew, Harenna shrew, Jenkins's shrew, Nicobar shrew, and Wimmer's shrew are categorized as critically endangered.
The 235 extant species of Crocidurinae are divided into nine genera; 192 of them are in Crocidura, the most species of any mammal genus.[2] A few extinct prehistoric Crocidurinae species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[3]
Conventions
| Conservation status | |
|---|---|
| EX | Extinct (0 species) |
| EW | Extinct in the wild (0 species) |
| CR | Critically Endangered (6 species) |
| EN | Endangered (21 species) |
| VU | Vulnerable (12 species) |
| NT | Near threatened (7 species) |
| LC | Least concern (128 species) |
| Other categories | |
| DD | Data deficient (60 species) |
| NE | Not evaluated (0 species) |
Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the crocidurine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.
Classification
The subfamily Crocidurinae consists of nine genera: Crocidura, containing 192 species; Diplomesodon and Feroculus with a single species each; Paracrocidura, with three species; Ruwenzorisorex, with one species; Scutisorex, containing two species; Solisorex, with one species, Suncus, containing nineteen species; and Sylvisorex, containing fifteen species.
Subfamily Crocidurinae
- Genus Crocidura (white-toothed shrews): 191 species
- Genus Diplomesodon (piebald shrew): one species
- Genus Feroculus (Kelaart's long-clawed shrew): one species
- Genus Paracrocidura (large-headed shrews): three species
- Genus Ruwenzorisorex (Ruwenzori shrew): one species
- Genus Scutisorex (hero shrews): two species
- Genus Solisorex (Pearson's long-clawed shrew): one species
- Genus Suncus (musk shrews and pygmy shrews): nineteen species
- Genus Sylvisorex (forest shrews): fifteen species
Crocidurines
The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[5]
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| African black shrew | C. nigrofusca Matschie, 1895 | Central and eastern Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[7] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| African dusky shrew | C. caliginea Hollister, 1916 | Central Africa | Size: 6–8 cm (2–3 in) long, plus 4–6 cm (2 in) tail[8] Habitat: Forest[9] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| African giant shrew
| C. olivieri (Lesson, 1827) 20 subspecies
| Sub-Saharan Africa | Size: 11–14 cm (4–6 in) long, plus 8–10 cm (3–4 in) tail[10] Habitat: Forest, savanna, and shrubland[11] Diet: Invertebrates[10] | LC
|
| Andaman shrew | C. andamanensis Miller, 1902 | South Andaman Island in India | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[12] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | CR
|
| Andaman spiny shrew | C. hispida Thomas, 1913 | South Andaman Island in India | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[13] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | VU
|
| Annamite shrew | C. annamitensis Jenkins, Lunde, & Moncrieff, 2009 | Vietnam | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[14] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Ansell's shrew | C. ansellorum Hutterer & Dippenaar, 1987 | South-central Africa | Size: 5–6 cm (2–2 in) long, plus 4–6 cm (2 in) tail[15] Habitat: Forest[16] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | EN
|
| Arabian shrew | C. arabica Hutterer, 1988 | Oman and Yemen | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Unknown[17] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Armenian shrew | C. armenica Gureev, 1963 | Armenia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Rocky areas[18] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Asian gray shrew | C. attenuata H. Milne-Edwards, 1872 | Eastern and southeastern Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest, shrubland, and grassland[19] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Asian lesser white-toothed shrew
| C. shantungensis Miller, 1901 Two subspecies
| Eastern Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Shrubland and grassland[20] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Bailey's shrew | C. baileyi Osgood, 1936 | Ethiopia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Grassland[21] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Bale shrew | C. bottegoides Hutterer & Yalden, 1990 | Ethiopia | Size: 4–6 cm (2 in) long, plus 4–5 cm (2 in) tail[22] Habitat: Grassland and forest[23] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | EN
|
| Banka shrew | C. vosmaeri Jentink, 1888 | Bangka Island in Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[24] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Batak shrew | C. batakorum Hutterer, 2007 | Philippines | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[25] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Bates's shrew | C. batesi (Dollman, 1915) | Western Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[26] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Beccari's shrew | C. beccarii Dobson, 1886 | Sumatra | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[27] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Beletta shrew | C. yaldeni Lavrenchenko, Voyta, & Hutterer, 2016 | Ethiopia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[28] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Bicolored musk shrew | C. fuscomurina (Heuglin, 1865) | Sub-Saharan Africa | Size: 4–8 cm (2–3 in) long, plus 2–5 cm (1–2 in) tail[29] Habitat: Savanna, grassland, and desert[30] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Bicolored shrew
| C. leucodon (Hermann, 1780) | Europe and western Asia | Size: 6–9 cm (2–4 in) long, plus 2–5 cm (1–2 in) tail[31] Habitat: Shrubland and grassland[32] Diet: Small mammals, frogs, toads, lizards, and invertebrates[31] | LC
|
| Black-footed shrew
| C. nigripes Miller & Hollister, 1921 Two subspecies
| Sulawesi island in Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[33] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Blackish white-toothed shrew | C. nigricans Bocage, 1889 | Angola | Size: 6–9 cm (2–4 in) long, plus 4–7 cm (2–3 in) tail[34] Habitat: Unknown[35] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Bornean shrew | C. foetida Peters, 1870 Three subspecies
| Borneo | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[36] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Bottego's shrew | C. bottegi Thomas, 1898 | Ethiopia | Size: 4–6 cm (2 in) long, plus 2–3 cm (1 in) tail[37] Habitat: Unknown[38] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Buettikofer's shrew | C. buettikoferi Jentink, 1888 | West-central Africa | Size: 7–10 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[39] Habitat: Forest[40] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Butiaba naked-tailed shrew | C. littoralis Heller, 1910 | Central Africa | Size: 8–10 cm (3–4 in) long, plus 6–8 cm (2–3 in) tail[41] Habitat: Forest[42] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Cameroonian shrew | C. picea Sanderson, 1940 | West-central Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest and grassland[43] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | EN
|
| Canarian shrew
| C. canariensis Hutterer, López-Jurado, & Vogel, 1987 | Canary Islands | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Coastal marine, desert, and shrubland[44] Diet: Snails and insects[44] | EN
|
| Caspian shrew | C. caspica Thomas, 1907 | Iran and Azerbaijan | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Unknown[45] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Chinese white-toothed shrew | C. rapax Allen, 1923 Four subspecies
| Eastern Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Unknown[46] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Christmas Island shrew | C. trichura Dobson, 1889 | Christmas Island | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[47] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | CR
|
| Cinderella shrew | C. cinderella Thomas, 1911 | Western Africa | Size: 5–7 cm (2–3 in) long, plus 4–5 cm (2–2 in) tail[48] Habitat: Savanna[49] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Congo white-toothed shrew | C. congobelgica Hollister, 1916 | Central Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[50] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Cranbrook's white-toothed shrew | C. cranbrooki Jenkins, Lunde, & Moncrieff, 2009 | Northern Myanmar | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[51] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Cretan shrew | C. zimmermanni Wettstein, 1953 | Island of Crete in Greece | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Unknown and rocky areas[52] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | EN
|
| Crosse's shrew | C. crossei Thomas, 1895 | Western Africa | Size: 6–7 cm (2–3 in) long, plus 4–6 cm (2 in) tail[53] Habitat: Forest[54] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Cyrenaica shrew | C. aleksandrisi Vesmanis, 1977 | Northern Libya | Size: 5–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[55] Habitat: Shrubland[56] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Dent's shrew
| C. denti Dollman, 1915 | Western and central Africa | Size: About 6 cm (2 in) long, plus 5 cm (2 in) tail[57] Habitat: Forest and savanna[58] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Desert musk shrew | C. smithii Thomas, 1895 Two subspecies
| Western and Eastern Africa | Size: 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[59] Habitat: Savanna[60] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Desperate shrew | C. desperata Hutterer, Jenkins, & Verheyen, 1991 | Tanzania | Size: 8–11 cm (3–4 in) long, plus about 7 cm (3 in) tail[61] Habitat: Forest[62] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | VU
|
| Dhofar shrew | C. dhofarensis (Hutterer, 1988) | Yemen and Oman | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Unknown[63] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Doucet's musk shrew | C. douceti Heim de Balsac, 1958 | Western Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[64] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Dsinezumi shrew
| C. dsinezumi (Temminck, 1842) | Japan | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Shrubland, grassland, and inland wetlands[65] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| East African highland shrew | C. allex Osgood, 1910 | Kenya and Tanzania | Size: 5–7 cm (2–3 in) long, plus 4–6 cm (2 in) tail[66] Habitat: Grassland and inland wetlands[67] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Egyptian pygmy shrew | C. religiosa (Geoffroy, 1827) | Northern Egypt | Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[68] Habitat: [69] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Eisentraut's shrew | C. eisentrauti Heim de Balsac, 1957 | Cameroon | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Grassland and forest[70] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | VU
|
| Elgon shrew | C. elgonius Osgood, 1910 | Kenya and Tanzania | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[71] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Elongated shrew | C. elongata Miller & Hollister, 1921 | Sulawesi island in Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest and shrubland[72] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Fingui white-toothed shrew | C. fingui Ceríaco et al., 2015 | Príncipe in São Tomé and Príncipe | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[73] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Fischer's shrew | C. fischeri Pagenstecher, 1885 | Kenya and Tanzania | Size: 6–10 cm (2–4 in) long, plus 4–6 cm (2 in) tail[74] Habitat: Savanna[75] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Flat-headed shrew | C. planiceps Heller, 1910 | West-central Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Inland wetlands[76] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Flower's shrew | C. floweri Dollman, 1915 | Northern Egypt | Size: 5–8 cm (2–3 in) long, plus 5–6 cm (2–2 in) tail[77] Habitat: Artificial terrain[78] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Fox's shrew | C. foxi Dollman, 1915 | Western and central Africa | Size: 9–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[79] Habitat: Savanna and forest[80] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Fraser's musk shrew | C. poensis (Fraser, 1843) | Western Africa | Size: 7–10 cm (3–4 in) long, plus 4–7 cm (2–3 in) tail[81] Habitat: Shrubland and forest[82] Diet: Arthropods and other invertebrates[81] | LC
|
| Gathorne's shrew | C. gathornei Jenkins, 2013 | Himalayas in India | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Shrubland and grassland[83] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Glass's shrew | C. glassi Heim de Balsac, 1966 | Ethiopia | Size: 6–9 cm (2–4 in) long, plus 4–6 cm (2 in)tail[84] Habitat: Shrubland, grassland, and inland wetlands[85] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | NT
|
| Gmelin's white-toothed shrew | C. gmelini Pallas, 1811 | Central Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Shrubland and grassland[86] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Goliath shrew | C. goliath Thomas, 1906 | Central Africa | Size: 15–18 cm (6–7 in) long, plus 10–14 cm (4–6 in) tail[87] Habitat: Forest[88] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Gracile naked-tailed shrew | C. maurisca Thomas, 1904 | Central Africa | Size: 7–10 cm (3–4 in) long, plus 6–8 cm (2–3 in) tail[89] Habitat: Inland wetlands and forest[90] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Grasse's shrew | C. grassei Brosset, Dubost, & Heim de Balsac, 1965 | West-central Africa | Size: 7–9 cm (3–4 in) long, plus 6–8 cm (2–3 in) tail[91] Habitat: Forest[92] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Greater Mindanao shrew | C. grandis Miller, 1911 | Philippines | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[93] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Greater red musk shrew
| C. flavescens (Geoffroy, 1827) | Southern Africa | Size: 8–12 cm (3–5 in) long, plus 3–6 cm (1–2 in) tail[94] Habitat: Savanna and grassland[95] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Greater white-toothed shrew
| C. russula (Hermann, 1780) | Europe and northwestern Africa | Size: 6–9 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[96] Habitat: Shrubland[97] Diet: Invertebrates, as well as small mammals and small lizards[96] | LC
|
| Greenwood's shrew | C. greenwoodi Heim de Balsac, 1966 | Somalia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest, savanna, and shrubland[98] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Guramba shrew | C. phaeura Osgood, 1936 | Ethiopia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[99] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | EN
|
| Guy's shrew | C. guy Jenkins, Lunde, & Moncrieff, 2009 | Vietnam | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[100] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Güldenstädt's shrew | C. gueldenstaedtii (Pallas, 1811) | Europe and western Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Shrubland, grassland, inland wetlands, rocky areas, intertidal marine, and coastal marine[101] Diet: Insects, as well as molluscs, amphipods, annelids, and small vertebrates[101] | LC
|
| Hainan Island shrew | C. wuchihensis Sung, 1966 | Southern China and northern Vietnam | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[102] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Harenna shrew | C. harenna Hutterer & Yalden, 1990 | Ethiopia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[103] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | CR
|
| Heather shrew | C. erica Dollman, 1915 | Angola | Size: About 10 cm (4 in) long, plus 5 cm (2 in) tail[104] Habitat: Unknown[105] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Hildegarde's shrew | C. hildegardeae Thomas, 1904 | Central Africa | Size: 6–8 cm (2–3 in) long, plus 3–6 cm (1–2 in) tail[106] Habitat: Forest[107] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Hill's shrew | C. hilliana Jenkins & Smith, 1995 | Thailand and Laos | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest and shrubland[108] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Horsfield's shrew | C. horsfieldii (Tomes, 1856) | Scattered southern Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[109] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Hun shrew | C. attila Dollman, 1915 | Central Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[110] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Hutan shrew | C. hutanis Ruedi & Vogel, 1995 | Sumatra island in Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[111] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Indochinese shrew | C. indochinensis Kloss, 1922 | Southeastern Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[112] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Iranian shrew | C. susiana Redding & Lay, 1978 | Iran | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Unknown[113] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Jackass shrew | C. arispa Spitzenberger, 1971 | Turkey | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Rocky areas[114] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Jackson's shrew | C. jacksoni Thomas, 1904 | East-central Africa | Size: 5–9 cm (2–4 in) long, plus 4–7 cm (2–3 in) tail[115] Habitat: Forest[116] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Javan ghost shrew | C. umbra Demos, Achmadi, Handika, Maharadatunkamsi, Rowe, & Esselstyn, 2016 | Java island in Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[117] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | NT
|
| Javanese shrew | C. maxi Sody, 1936 | Indonesia and East Timor | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[118] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Jenkins's shrew | C. jenkinsi Chakraborty, 1978 | South Andaman Island in India | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[119] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | CR
|
| Jouvenet's shrew | C. jouvenetae Heim de Balsac, 1958 | Western Africa | Size: 6–9 cm (2–4 in) long, plus 4–7 cm (2–3 in) tail[120] Habitat: Forest[121] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Kashmir white-toothed shrew | C. pullata Miller, 1911 | Northern India and Pakistan | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Unknown[122] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Katinka's shrew | C. katinka Bate, 1937 | Western Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Unknown[123] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Kilimanjaro shrew | C. monax Thomas, 1910 |  | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[124] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Kinabalu shrew | C. baluensis Thomas, 1898 | Northern Borneo | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest, shrubland, and grassland[125] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Kivu long-haired shrew | C. lanosa Heim de Balsac, 1968 | Central Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest and inland wetlands[126] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | VU
|
| Kivu shrew | C. kivuana Heim de Balsac, 1968 | Central Africa | Size: 6–9 cm (2–4 in) long, plus 6–8 cm (2–3 in) tail[127] Habitat: Forest and inland wetlands[128] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | NT
|
| Lamotte's shrew | C. lamottei Heim de Balsac, 1968 | Western Africa | Size: 8–9 cm (3–4 in) long, plus 3–6 cm (1–2 in) tail[127] Habitat: Forest and savanna[129] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Large-headed forest shrew | C. grandiceps Hutterer, 1983 | Western Africa | Size: 9–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[130] Habitat: Forest[131] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Latona's shrew | C. latona Hollister, 1916 | Democratic Republic of the Congo | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[132] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Lesser Ryukyu shrew | C. watasei Kuroda, 1924 | Southern Japan | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Shrubland, grassland, and inland wetlands[133] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Lesser gray-brown musk shrew | C. silacea Thomas, 1895 | Southern Africa | Size: 6–8 cm (2–3 in) long, plus 4–6 cm (2 in) tail[134] Habitat: Rocky areas, grassland, shrubland, and savanna[135] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Lesser red musk shrew
| C. hirta Peters, 1852 | Southern and southeastern Africa | Size: 6–10 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[136] Habitat: Savanna, grassland, and desert[137] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Lesser rock shrew | C. serezkyensis Laptev, 1929 | Western Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Unknown[138] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Lesser white-toothed shrew
| C. suaveolens (Pallas, 1811) | Europe and Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest, shrubland, grassland, rocky areas, and coastal/supratidal marine[139] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Long-footed shrew | C. crenata Brosset, Dubost, & Heim de Balsac, 1965 | West-central Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[140] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Long-tailed musk shrew | C. dolichura Peters, 1876 | Central Africa | Size: 6–7 cm (2–3 in) long, plus 7–8 cm (3 in) tail[141] Habitat: Forest[142] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Lucina's shrew | C. lucina Dippenaar, 1980 | Ethiopia | Size: 8–10 cm (3–4 in) long, plus 4–6 cm (2 in) tail[143] Habitat: Grassland and inland wetlands[144] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | VU
|
| Ludia's shrew | C. ludia Hollister, 1916 | Central Africa | Size: 6–7 cm (2–3 in) long, plus 5–6 cm (2–2 in) tail[145] Habitat: Forest[146] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Luzon shrew | C. grayi Dobson, 1890 | Philippines | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[147] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Lwiro shrew | C. lwiroensis Peterhans & Hutterer, 2013 | Democratic Republic of the Congo | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[148] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| MacArthur's shrew | C. macarthuri St. Leger, 1934 | Eastern Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Savanna[149] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| MacMillan's shrew | C. macmillani Dollman, 1915 | Ethiopia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest and savanna[150] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | NT
|
| Makwassie musk shrew | C. maquassiensis Roberts, 1946 | Southern Africa | Size: 5–8 cm (2–3 in) long, plus 4–5 cm (2–2 in) tail[151] Habitat: Grassland and rocky areas[152] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Malayan shrew | C. malayana Robinson & Kloss, 1911 | Malaysia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[153] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Mamfe shrew | C. virgata Sanderson, 1940 | West-central Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest and grassland[154] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Manenguba shrew | C. manengubae Hutterer, 1982 | West-central Africa | Size: 7–9 cm (3–4 in) long, plus 6–7 cm (2–3 in) tail[151] Habitat: Forest, shrubland, and grassland[155] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | VU
|
| Mauritanian shrew | C. lusitania Dollman, 1915 | Western and eastern Africa | Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[156] Habitat: Savanna and shrubland[157] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Mduma's shrew | C. mdumai Stanley, Hutterer, Giarla, & Esselstyn, 2015 | Tanzania | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[158] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | EN
|
| Mindanao shrew
| C. beatus Miller, 1910 | Philippines | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[159] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Mindoro shrew | C. mindorus Miller, 1910 | Philippines | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[160] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Montane white-toothed shrew | C. montis Thomas, 1906 | Eastern Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest, shrubland, and grassland[161] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Moonshine shrew | C. luna Dollman, 1910 | Southeastern Africa | Size: 6–11 cm (2–4 in) long, plus 3–7 cm (1–3 in) tail[156] Habitat: Forest[162] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Mossy forest shrew | C. musseri Ruedi & Vogel, 1995 | Sulawesi island in Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[163] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Mount Nimba giant forest shrew | C. nimbasilvanus Hutterer, 2003 | Western Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[164] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Munissi's shrew | C. munissii Stanley, Hutterer, Giarla, & Esselstyn, 2015 | Tanzania | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[165] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Narrow-headed shrew | C. stenocephala Heim de Balsac & Dieterlen, 1979 | Central Africa | Size: 8–11 cm (3–4 in) long, plus 6–9 cm (2–4 in) tail[166] Habitat: Inland wetlands[167] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | VU
|
| Negev shrew | C. ramona Ivanitskaya, Shenbrot, & Nevo, 1996 | Israel | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Desert[168] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Negros shrew | C. negrina Rabor, 1952 | Philippines | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[169] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | EN
|
| Nicobar shrew | C. nicobarica Miller, 1902 | Great Nicobar Island in India | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[170] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | CR
|
| Nigerian shrew | C. nigeriae Dollman, 1915 | West-central Africa | Size: 9–11 cm (4 in) long, plus 5–7 cm (2–3 in) tail[171] Habitat: Forest[172] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Nimba shrew | C. nimbae Heim de Balsac, 1956 | Western Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[173] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | NT
|
| Niobe's shrew | C. niobe Thomas, 1906 | Central Africa | Size: 6–9 cm (2–4 in) long, plus 5–7 cm (2–3 in) tail[174] Habitat: Forest and grassland[175] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| North African white-toothed shrew
| C. pachyura Küster, 1835 | Northern Africa and southern Europe | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Shrubland[176] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Nyiro shrew | C. macowi Dollman, 1915 | Kenya | Size: 6–8 cm (2–3 in) long, plus 5–6 cm (2–2 in) tail[177] Habitat: Unknown[178] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Oriental shrew
| C. orientalis Jentink, 1890 Two subspecies
| Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[179] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Palawan shrew | C. palawanensis Taylor, 1934 | Philippines | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest and shrubland[180] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Pale gray shrew | C. pergrisea Miller, 1913 | Pakistan | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[181] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Panay shrew | C. panayensis Hutterer, 2007 | Philippines | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[182] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Peninsular shrew | C. negligens Kloss, 1914 | Southeastern Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[183] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Peters's musk shrew | C. gracilipes Peters, 1870 | Tanzania | Size: About 6 cm (2 in) long, plus 5 cm (2 in) tail[87] Habitat: Unknown[184] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Phan Luong's shrew | C. phanluongi Jenkins, Abramov, Rozhnov, & Olsson, 2010 | Vietnam and Cambodia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[185] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Phu Hoc shrew | C. phuquocensis Abramov, Jenkins, Rozhnov, & Kalinin, 2008 | Phú Quốc island in Vietnam | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[186] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Pitman's shrew | C. pitmani Barclay, 1932 | Zambia | Size: About 5 cm (2 in) long, plus 4 cm (2 in) tail[187] Habitat: Unknown[188] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Polia's shrew | C. polia Hollister, 1916 | Democratic Republic of the Congo | Size: About 6 cm (2 in) long, plus 7 cm (3 in) tail[81] Habitat: Unknown[189] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Rainey's shrew | C. raineyi Heller, 1912 | Kenya | Size: 9–10 cm (4 in) long, plus 5–7 cm (2–3 in) tail[190] Habitat: Forest[191] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Reddish-gray musk shrew | C. cyanea (Duvernoy, 1838) | Southern Africa | Size: 6–10 cm (2–4 in) long, plus 3–7 cm (1–3 in) tail[192] Habitat: Forest, grassland, and caves[193] Diet: Invertebrates[193] | LC
|
| Roosevelt's shrew | C. roosevelti (Heller, 1910) | Central Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Savanna[194] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Ryukyu shrew | C. orii Kuroda, 1924 | Southern Japan | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[195] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | EN
|
| Sa Pa shrew
| C. sapaensis Jenkins, Abramov, Bannikova, & Rozhnov, 2013 | Vietnam | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest and grassland[196] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Saharan shrew | C. tarfayensis Vesmanis & Vesmanis, 1980 | Mauritania and Morocco | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Rocky areas and desert[197] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Sahelian tiny shrew | C. pasha Dollman, 1915 | Scattered northern Africa | Size: 4–6 cm (2 in) long, plus 2–4 cm (1–2 in) tail[198] Habitat: Savanna[199] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Savanna dwarf shrew | C. nanilla Thomas, 1909 | Western and eastern Africa | Size: 4–6 cm (2–2 in) long, plus 3–5 cm (1–2 in) tail[200] Habitat: Savanna[201] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Savanna path shrew | C. viaria (Geoffroy, 1834) | Western, central, and eastern Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest and savanna[202] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Savanna shrew | C. fulvastra (Sundevall, 1843) | Central Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Savanna[203] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Savanna swamp shrew | C. longipes Hutterer & Happold, 1983 | West-central Africa | Size: 9–12 cm (4–5 in) long, plus 5–7 cm (2–3 in) tail[204] Habitat: Inland wetlands[205] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Siberian shrew | C. sibirica Dukelsky, 1930 | Central Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest, grassland, inland wetlands, and desert[206] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Sibuyan shrew | C. ninoyi Esselstyn & Goodman, 2010 | Philippines | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[207] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Sicilian shrew
| C. sicula Miller, 1900 Four subspecies
| Italy and Malta | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Shrubland[208] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Sinharaja white-toothed shrew | C. hikmiya Meegaskumbura, Meegaskumbura, Pethiyagoda, Manamendra-Arachchi, & Schneider, 2007 | Sri Lanka | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[209] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | EN
|
| Small-footed shrew | C. parvipes Osgood, 1910 | Central Africa | Size: 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[210] Habitat: Forest, savanna, and shrubland[211] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Smoky white-toothed shrew | C. fumosa Thomas, 1904 | Kenya | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[212] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Sokolov's shrew | C. sokolovi Jenkins, Abramov, Rozhnov, & Makarova, 2007 | Vietnam | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[213] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Somali dwarf shrew | C. nana Dobson, 1890 | Ethiopia and Somalia | Size: About 4 cm (2 in) long, plus 3 cm (1 in) tail[214] Habitat: Grassland[215] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Somali shrew | C. somalica Thomas, 1895 | Eastern Africa | Size: 6–8 cm (2–3 in) long, plus 4–5 cm (2–2 in) tail[166] Habitat: Savanna[216] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Southeast Asian shrew
| C. fuliginosa (Blyth, 1856) | Southeastern Asian | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest, shrubland, grassland, and rocky areas[217] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Sri Lankan long-tailed shrew | C. miya Phillips, 1929 | Sri Lanka | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[218] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | EN
|
| Sulawesi shrew | C. lea Miller & Hollister, 1921 | Sulawesi island in Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[219] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Sulawesi tiny shrew | C. levicula Miller & Hollister, 1921 | Sulawesi island in Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[220] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Sulawesi white-handed shrew | C. rhoditis Miller & Hollister, 1921 | Sulawesi island in Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[221] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Sumatran giant shrew | C. lepidura Lyon, 1908 | Sumatra island in Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest and shrubland[222] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Sumatran long-tailed shrew | C. paradoxura Dobson, 1886 | Sumatra island in Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[223] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Sunda shrew
| C. monticola Peters, 1870 | Southeastern Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[224] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Swamp musk shrew | C. mariquensis (Smith, 1844) Three subspecies
| Southern Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Inland wetlands[225] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| São Tomé shrew
| C. thomensis (Bocage, 1887) | São Tomé Island in São Tomé and Príncipe | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[226] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | EN
|
| Taiwanese gray shrew | C. tanakae Kuroda, 1938 | Eastern Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest and grassland[227] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Tanzanian shrew | C. tansaniana Hutterer, 1986 | Tanzania | Size: 8–11 cm (3–4 in) long, plus 5–8 cm (2–3 in) tail[228] Habitat: Forest[229] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | NT
|
| Tarella shrew | C. tarella Dollman, 1915 | Central Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[230] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | EN
|
| Telford's shrew | C. telfordi Hutterer, 1986 | Tanzania | Size: 10–11 cm (4 in) long, plus 7–9 cm (3–4 in) tail[231] Habitat: Forest[232] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | VU
|
| Thalia's shrew | C. thalia Dippenaar, 1980 | Ethiopia | Size: 7–10 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[231] Habitat: Forest, savanna, shrubland, grassland, and inland wetlands[233] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Therese's shrew | C. theresae Heim de Balsac, 1968 | Western Africa | Size: 7–11 cm (3–4 in) long, plus 3–6 cm (1–2 in) tail[234] Habitat: Forest and savanna[235] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Thick-tailed shrew | C. brunnea Jentink, 1888 Two subspecies
| Indonesia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[236] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Timor shrew | C. tenuis (Müller, 1840) | Timor | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[237] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Turbo shrew | C. turba Dollman, 1910 | Central Africa | Size: 9–11 cm (4 in) long, plus 5–6 cm (2–2 in) tail[238] Habitat: Forest[239] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Ugandan lowland shrew | C. selina (Dollman, 1915) | Kenya and Uganda | Size: 7–9 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[240] Habitat: Forest and inland wetlands[241] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Ugandan musk shrew | C. mutesae Heller, 1910 | Central Africa | Size: About 11 cm (4 in) long, plus 6 cm (2 in) tail[214] Habitat: Forest[242] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Ultimate shrew | C. ultima Dollman, 1915 | Kenya | Size: About 9 cm (4 in) long, plus 6 cm (2 in) tail[243] Habitat: Forest[244] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Upemba shrew | C. zimmeri Osgood, 1936 | Democratic Republic of the Congo | Size: 11–13 cm (4–5 in) long, plus 4–6 cm (2–2 in) tail[245] Habitat: Unknown[246] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Usambara shrew | C. usambarae Dippenaar, 1980 | Tanzania | Size: 7–9 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[247] Habitat: Forest[248] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | VU
|
| Ussuri white-toothed shrew
| C. lasiura (Dobson, 1890) | Eastern Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Grassland[249] Diet: Insects and other invertebrates, as well as small vertebrates[249] | LC
|
| Voi shrew | C. voi Osgood, 1910 | Central and eastern Africa | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Savanna[250] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Voracious shrew
| C. vorax Allen, 1923 | Southeastern Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Forest[251] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| West African long-tailed shrew | C. muricauda (Miller, 1900) | Western Africa | Size: 5–7 cm (2–3 in) long, plus tail[252] Habitat: Forest[253] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| West African pygmy shrew | C. obscurior Heim de Balsac, 1958 | Western Africa | Size: 4–5 cm (2–2 in) long, plus 3–4 cm (1–2 in) tail[254] Habitat: Forest[255] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Whitaker's shrew | C. whitakeri De Winton, 1898 | Northwestern Africa | Size: 5–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[256] Habitat: Coastal marine, intertidal marine, and shrubland[257] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Wimmer's shrew | C. wimmeri Heim de Balsac & Aellen, 1958 | Ivory Coast | Size: 8–9 cm (3–4 in) long, plus 5–6 cm (2–2 in) tail[258] Habitat: Forest[259] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | CR
|
| Xanthippe's shrew | C. xantippe Osgood, 1910 | Kenya and Tanzania | Size: 8–10 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[260] Habitat: Savanna and shrubland[261] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Yankari shrew | C. yankariensis Hutterer & Jenkins, 1980 | Central and eastern Africa | Size: 5–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[260] Habitat: Savanna[262] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
| Zaphir's shrew | C. zaphiri Dollman, 1915 | Ethiopia and Kenya | Size: About 10 cm (4 in) long, plus 6 cm (2 in) tail[263] Habitat: Unknown[264] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | DD
|
| Zarudny's rock shrew
| C. zarudnyi Ognew, 1928 | Central Asia | Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6] Habitat: Desert[265] Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] | LC
|
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| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Piebald shrew
| D. pulchellum (Lichtenstein, 1823) | Central Asia | Size: 5–8 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[266] Habitat: Desert[267] Diet: Insects, as well as small lizards[267] | LC
|
,_Gescheckte_W%C3%BCstenspitzmaus,_Piebald_Shrew,_%D0%9C%D0%BE%D1%81%D0%BA%D0%BE%D0%B2%D1%81%D0%BA%D0%B8%D0%B9_%D0%B7%D0%BE%D0%BE%D0%BF%D0%B0%D1%80%D0%BA_(Moscow_zoo),_18.11.2008.jpg)
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Kelaart's long-clawed shrew | F. feroculus (Kelaart, 1850) | Southern India and Sri Lanka | Size: 10–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[268] Habitat: Forest, grassland, and inland wetlands[269] Diet: Earthworms, small vertebrates, and plants[268] | EN
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Grauer's large-headed shrew | P. graueri Hutterer, 1986 | Democratic Republic of the Congo | Size: 6–10 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[270] Habitat: Forest[271] Diet: Insects[270] | DD
|
| Greater large-headed shrew | P. maxima Heim de Balsac, 1959 | Central Africa | Size: 6–10 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[270] Habitat: Forest and inland wetlands[272] Diet: Insects[270] | LC
|
| Lesser large-headed shrew | P. schoutedeni Heim de Balsac, 1956 | West-central Africa | Size: 6–10 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[270] Habitat: Forest[273] Diet: Insects[270] | LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Ruwenzori shrew | R. suncoides (Osgood, 1936) | Central Africa | Size: 9–10 cm (4 in) long, plus 5–7 cm (2–3 in) tail[274] Habitat: Inland wetlands and forest[275] Diet: Mollusks and small vertebrates[268] | VU
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Hero shrew
| S. somereni (Thomas, 1910) | Central Africa | Size: 12–15 cm (5–6 in) long, plus 6–10 cm (2–4 in) tail[276] Habitat: Forest[277] Diet: Insects, earthworms, and small vertebrates[276] | LC
|
| Thor's hero shrew | S. thori Stanley, Malekani, &Gambalemoke, 2013 | Democratic Republic of the Congo | Size: 12–15 cm (5–6 in) long, plus 6–10 cm (2–4 in) tail[278] Habitat: Forest[279] Diet: Small animals and plants[278] | DD
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Pearson's long-clawed shrew | S. pearsoni Thomas, 1924 | Sri Lanka | Size: 12–15 cm (5–6 in) long, plus 5–7 cm (2–3 in) tail[266] Habitat: Forest and grassland[280] Diet: Insects, earthworms, and small vertebrates[266] | EN
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Anderson's shrew | S. stoliczkanus (Anderson, 1877) | Southern Asia | Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281] Habitat: Forest and grassland[282] Diet: Insects[281] | LC
|
| Asian highland shrew | S. montanus (Kelaart, 1850) | Southern India and Sri Lanka | Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281] Habitat: Forest and grassland[283] Diet: Insects[281] | LC
|
| Asian house shrew
| S. murinus (Linnaeus, 1766) | Asia and Africa (introduced in red) | Size: 10–5 cm (4–2 in) long, plus 5–9 cm (2–4 in) tail[284] Habitat: Forest, savanna, shrubland, grassland, and inland wetlands[285] Diet: Insects and small mammals, as well as plants[286] | LC
|
| Black shrew | S. ater (Medway, 1965) | Northern Borneo | Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281] Habitat: Forest[287] Diet: Insects[281] | DD
|
| Bornean pygmy shrew | S. hosei (Thomas, 1893) | Borneo | Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281] Habitat: Forest[288] Diet: Insects[281] | DD
|
| Climbing shrew | S. megalura (Jentink, 1888) | Sub-Saharan Africa | Size: 5–6 cm (2 in) long, plus 8–10 cm (3–4 in) tail[284] Habitat: Forest and savanna[289] Diet: Insects[284] | LC
|
| Day's shrew | S. dayi (Dodson, 1888) | Southern India | Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281] Habitat: Forest and grassland[290] Diet: Insects[281] | EN
|
| Etruscan shrew
| S. etruscus (Savi, 1822) | Europe, northern Africa, and Asia | Size: 3–5 cm (1–2 in) long, plus 2–3 cm (1 in) tail[1] Habitat: Forest, shrubland, grassland, and desert[291] Diet: Insects[292] | LC
|
| Flores shrew | S. mertensi Kock, 1974 | Flores island in Indonesia | Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281] Habitat: Forest[293] Diet: Insects[281] | EN
|
| Greater dwarf shrew
| S. lixus (Thomas, 1898) | Southern Africa | Size: 5–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[281] Habitat: Forest, savanna, shrubland, and grassland[294] Diet: Insects[281] | LC
|
| Hutu-Tutsi dwarf shrew | S. hututsi Peterhans & Hutterer, 2009 | Central Africa | Size: 4–6 cm (2 in) long, plus 2–3 cm (1 in) tail[281] Habitat: Forest and shrubland[295] Diet: Insects[281] | DD
|
| Jungle shrew | S. zeylanicus Phillips, 1928 | Sri Lanka | Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281] Habitat: Forest[296] Diet: Insects[281] | EN
|
| Least dwarf shrew | S. infinitesimus (Heller, 1912) | South Africa and Kenya | Size: 5–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[281] Habitat: Forest and grassland[297] Diet: Insects[281] | LC
|
| Lesser dwarf shrew | S. varilla (Thomas, 1895) | Southern Africa | Size: 5–7 cm (2–3 in) long, plus 2–5 cm (1–2 in) tail[281] Habitat: Forest, savanna, shrubland, and grassland[298] Diet: Insects[281] | LC
|
| Malayan pygmy shrew | S. malayanus (Kloss, 1917) | Malaysia and Thailand | Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281] Habitat: Forest[299] Diet: Insects[281] | LC
|
| Remy's pygmy shrew | S. remyi Brosset, Dubost, & Heim de Balsac, 1965 | West-central Africa | Size: 4–5 cm (2 in) long, plus 1–3 cm (0.4–1.2 in) tail[281] Habitat: Forest[300] Diet: Insects[281] | LC
|
| Sri Lankan shrew | S. fellowesgordoni Phillips, 1932 | Sri Lanka | Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281] Habitat: Forest and grassland[301] Diet: Insects[281] | EN
|
| Taita shrew | S. aequatorius (Heller, 1912) | Kenya | Size: 6–9 cm (2–4 in) long, plus 5–7 cm (2–3 in) tail[302] Habitat: Forest[303] Diet: Insects[304] | EN
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Bamenda pygmy shrew | S. silvanorum (Hutterer, Riegert, & Sedláček, 2009) | Cameroon | Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[274] Habitat: Forest and grassland[305] Diet: Insects[306] | DD
|
| Bioko forest shrew | S. isabellae Heim de Balsac, 1968 | West-central Africa | Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[274] Habitat: Forest and grassland[307] Diet: Insects[306] | VU
|
| Cameroonian forest shrew | S. camerunensis Heim de Balsac, 1968 | West-central Africa | Size: 5–6 cm (2 in) long, plus 5–7 cm (2–3 in) tail[308] Habitat: Forest and grassland[309] Diet: Insects[306] | VU
|
| Corbet's forest shrew | S. corbeti Hutterer, 2009 | West-central Africa | Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[274] Habitat: Forest[310] Diet: Insects[306] | DD
|
| Dudu Akaibe's pygmy shrew | S. akaibei Mukinzi, Hutterer, & Barriere, 2009 | Democratic Republic of the Congo | Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[274] Habitat: Forest[311] Diet: Insects[306] | DD
|
| Grant's forest shrew | S. granti Thomas, 1907 Two subspecies
| Central Africa | Size: 4–7 cm (2–3 in) long, plus 4–7 cm (2–3 in) tail[312] Habitat: Forest[313] Diet: Insects[306] | LC
|
| Greater forest shrew | S. ollula Thomas, 1913 | Central Africa | Size: 5–8 cm (2–3 in) long, plus 1–2 cm (0–1 in) tail[314] Habitat: Forest[315] Diet: Insects[306] | LC
|
| Howell's forest shrew | S. howelli Jenkins, 1984 | Tanzania | Size: 5–8 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[316] Habitat: Forest[317] Diet: Insects[306] | LC
|
| Johnston's forest shrew | S. johnstoni (Dobson, 1888) | West-central and central Africa | Size: 7–10 cm (3–4 in) long, plus 4–7 cm (2–3 in) tail[308] Habitat: Forest[318] Diet: Insects[306] | LC
|
| Kongana shrew | S. konganensis Ray & Hutterer, 1996 | Central Africa | Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[274] Habitat: Forest[319] Diet: Insects[306] | DD
|
| Lesser forest shrew
| S. oriundus Hollister, 1916 | Central Africa | Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[274] Habitat: Forest[320] Diet: Insects[306] | DD
|
| Moon forest shrew | S. lunaris Thomas, 1906 | Central Africa | Size: 7–10 cm (3–4 in) long, plus 4–6 cm (2 in) tail[321] Habitat: Forest[322] Diet: Insects[306] | NT
|
| Mount Cameroon forest shrew | S. morio (Gray, 1862) | West-central Africa | Size: 6–8 cm (2–3 in) long, plus 5–6 cm (2 in) tail[323] Habitat: Forest[324] Diet: Insects[306] | EN
|
| Rain forest shrew | S. pluvialis Hutterer & Schlitter, 1996 | West-central Africa | Size: 6–8 cm (2–3 in) long, plus 5–8 cm (2–3 in) tail[321] Habitat: Forest[325] Diet: Insects[306] | DD
|
| Volcano shrew | S. vulcanorum Hutterer & Verheyan, 1985 | Central Africa | Size: 4–6 cm (2 in) long, plus 4–7 cm (2–3 in) tail[308] Habitat: Forest and inland wetlands[326] Diet: Insects[306] | LC
|
References
Sources
- Kingdon, Jonathan (2014). Mammals of Africa. Vol. IV: Hedgehogs, Shrews and Bats. A & C Black. ISBN 978-1-4081-8993-1.
- Nowak, Ronald M. (1999). Walker's Mammals of the World. Vol. 1. Johns Hopkins University Press. ISBN 978-0-8018-5789-8.
- Hutterer, Rainer (2005). Wilson, Don E.; Reeder, DeeAnn M. (eds.). Mammal Species of the World. Vol. 1 (3rd ed.). Johns Hopkins University Press. ISBN 978-0-8018-8221-0.
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